Pulsatilla orientali-sibirica

Pulsatilla orientali-sibirica Stepanov sp. nov. – P. flavescens (Zucc.) Juz., 1937, Фл. СССР, 7: 296, p.p. non P. flavescens (Hazsl.) Borb. emend. Boros, 1924, Bot. Kozlem., 21(1–6): 65; Черепнин, Фл. юж. части Краснояр. кр., 1961, 3: 166; Положий, Ревердатто, 1976, Фл. Краснояр. кр., 5(3): 77; Степанов, 1996, Фл. анализ: 71; Штаркер, 1988, Тр. госзап. «Столбы», 15: 42; Штаркер, 1989, Тр. госзап. «Столбы», 16: 28; Сонникова, 1992, Сосуд. раст. Саяно-Шуш. заповедн.: 48; Тимохина, 1993, Фл. Сиб., 6:153; Андреева и др., 2010, Тр. гос. зап. «Столбы», 18: 100; Пешкова, 2001, Флороген. анализ степн. фл. Юж. Сиб.: 37. – P. uralensis auct.: Эбель, 2012, Конспект фл. северо-западной части Алт.-Саян. провинции: 129. – P. patens var. flavescens (Zucc.) Zämelis. – Крылов, 1931, Фл. Зап. Сиб., 5: 1165. – P. patens auct.: Верещагин, 1940, Инвент. фл. зап. «Столбы»: 36, р.р.; Антипова, 2012, Фл. внутриконт. островн. лесост. Ср. Сиб.: 138; Сонникова, 2012, Сосуд. раст. нац. парка «Шуш. бор»: 58; Шауло, 2006, Фл. Зап. Саяна: 44, р.р.; Луферов, 2008, Консп. фл. Иркут. обл.: 129; Попов, 1957, Фл. Ср. Сиб., 1: 247, р.р.; Пешкова, 1979, Фл. Центр. Сиб.: 359, р.р. – Anemone flavescens Zucc., 1826, Regensb. Zeit., 1: 371, non A. flavescens Rapaics, 1916.
Caules 5–30 (50) cm alt. Folia radicalia, hysteranthia, longipetiolata, laminis plus minusve rotundatis, digitatim sectis, trilaciniatis; laciniae laminarum sessiles, 2–3-dissectae lacinis dentatis ordinis secundi (in numero 18–40) 2–4 mm lat. Flores pallide-flavi, campanulati, interdum late aperti. Tepala 2.5–3.5 cm lg., 1.3–1.7 cm lat., ovata vel oblongo-ovata. Fructiculi pilosi aristis plumosis 2.5–3.5 cm lg. (Figure 1, B).
Typus: In adjacentibus urbis Krasnojarsk, collis Nikolaevskaja Sopka, declive stepposum. 25.04.2014. N.V. Stepanov (Holotypus – KRSU; isotypi – TK, LE).
Paratypi: Prov. Krasnojarskij, distr. Ermakovskij, viridarium nationale «Ergaki», vallis fl. Us prope stationem Idzhim, steppa petrosa. 05.05.2011. N.V. Stepanov (KRSU, TK); Prope urbem Krasnojarsk, ad finis borealem reservati ―Stolby‖, vallis fl. Laletina, declive austro-occidentale, steppa petrosa. 27.04.2014. N.V. Stepanov (KRSU). Urbs Krasnojarsk, prope Akademgorodok, declive stepposum, petrosum. 25.04.2013. N.V. Stepanov (KRSU); Urbs Krasnojarsk, collis Nikolaevskaja Sopka, declive stepposum, petrosum. 23.04.2013. N.V. Stepanov (KRSU); Urbs Krasnojarsk, area ripae fl. Enissej, pratum substepposum. 27.07.2001. E. Kostjutschenko (KRSU); Urbs Krasnojarsk, prope Akademgorodok, trames Monastyrskaja Tropa, steppa petrosa. 16.05.1999. N.V. Stepanov (KRSU); Urbs Krasnojarsk, collis Nikolaevskaja Sopka, declive australe. 20.05.1955. Vytschuzhanin, Ponomarenko (KRSU).
Affinitas. A Pulsatilla patenti s.str. floribus pallide-flavibus, minus apertibus, tepalis latis et abrupte acutatis vel obtusiusculis; a Pulsatilla uralensi (Zämelis) Tzvelev laciniis angustis, segmentis sessilibus; a Pulsatilla angustifolia Turcz. laciniis latis et oligomeris, segmentis sessilibus, evolutione foliis serotina; a speciebus commemoratis area (ab Altai in parte occidentale ad Transbaikal in parte orientale et ab Tuva in parte australe ad latitude fl. Nizhnjaja Tunguska in parte boreale) et inclinate ad habitationes stepposas differt.
Stems 5–30 (50) cm tall. Basal leaves develop at the end of flowering, long-petiolate, with lamina more or less rounded, palmately dissected into three sessile share; parts of lamina are twice-thrice dissected into dentate lobes of the second order (in number 18–40) 2–4 mm wide. Flowers are pale yellow, bell-shaped, sometimes broadly disclosed. Tepals (2.5–3.5) cm long, (1.3–1.7) cm wide, ovate or oblong-ovate. Fruitlets pubescent with long pinnate aristae 2.5–3.5 cm long (Figure 1, B).
Type: The vicinity of Krasnoyarsk, Nikolaevskaya Sopka hill, the southern slope. 25.04.2014. N.V. Stepanov (Holotype – KRSU); isotypes – TK, LE).
Paratypes: Krasnoyarskiy krai, Yermakovskiy district, the natural park "Ergaki", valley of the Us river, in areas of station Idzhim, rocky steppe. 05.05.2011. N.V. Stepanov (KRSU, TK); Env. Krasnoyarsk, near the northern boundary of the reserve "Stolby", valley of Laletina river, southwestern slope, rocky steppe. 27.04.2014. N.V. Stepanov (KRSU); Krasnoyarsk, rocky steppe slopes in Akademgorodok. 25.04.2013. N.V. Stepanov (KRSU); The vicinity of Krasnoyarsk, Nikolaevskaya Sopka hill, the southern slope. 04.23.2013. N.V. Stepanov (KRSU); Krasnoyarsk, exposed waterside areas of the Yenisei, steppefied meadow. 27.07.2001. E. Kostyuchenko (KRSU); Krasnoyarsk, Akademgorodok, Monastyrskaya trail, rocky steppe. 16.05.1999. N.V. Stepanov (KRSU); The vicinity of Krasnoyarsk, Nikolaevskaya Sopka hill, the southern slope. 20.05.1955. Vychuzhanin, Ponomarenko (KRAS, KRSU); 
Relationship. This taxon differs from closely related species Pulsatilla patens s.str. by yellowish, less open flower, tepals broader and abruptly pointed or obtuse; from Pulsatilla uralensis (Zämelis) Tzvelev by narrower segments sessile leaves; from Pulsatilla angustifolia Turcz. by wider and not numerous, sessile leaf lobes, later development of leaves; from all these species by the range (from the Altai in the west to the Transbaikalia in the east and from Tuva in the south to the latitude of NizhnyayaTunguska river in the north) and attraction to steppe habitats.   
p. ovczinnikovii Maximova